A stronger and more precise statement about the relationship between the wheel motions and the rotation of the chassis is “The total rotation of the chassis is proportional to the difference between the forwards-minus-backwards motions of the two wheels”.
If we take the rolling velocity of the left and right wheels as $\dot{\alpha}_{1}$ and $\dot{\alpha}_{2}$, the wheel radius as $R$ and the length of the axle as $L$, the turning velocity of the axle is
$\dot{\theta} = (-\dot{\alpha}_{1} + \dot{\alpha}_{2})\frac{R}{L}$.
Factoring out the time portion of the derivative makes infinitesimal changes in $\theta$ depend directly on infinitesimal changes in the roll angles $\alpha$,
$d\theta = (-d\alpha_{1} +d\alpha_{2})\frac{R}{L}$,
such that we can express the derivative of the axle orientation with respect to the wheel roll angles as the one-form $A$
$\frac{\partial \theta}{\partial \alpha} = \overbrace{\begin{bmatrix} -1 & 1 \end{bmatrix}}^{A}\frac{R}{L}$.
This one-form is constant, such that its exterior derivative (i.e. curl) is zero. This means that orientation is conservative with respect to the roll angles of the wheels: once the vehicle has been placed on the ground, the total change in orientation is a function of the net angles that the wheels have rolled through, and independent of the intermediate roll angles they passed through to get there. Here, specifically, the orientation relative to the starting orientation is proportional to the difference between the net angle each wheel has rolled through,
$\Delta\theta = (-\Delta\alpha_{1} + \Delta\alpha_{2})\frac{R}{L}$
The net forward-minus-backward motion of each wheel (measured at its hub) is directly proportional to the net roll angle of the wheel around its axle,
$n = R\Delta\alpha$,
so the orientation can also be expressed in terms of the net forwards-minus-backwards motion of each wheel,
$\Delta\theta = (-n_{1} + n_{2})/L$.
This measure of rotation change of the chassis distinguishes between returning to the original orientation and making a full rotation, so we can say that for a system at $\Delta\theta=0$, the net forward-minus-backward motions of the two wheels are equal.
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As noted in https://math.stackexchange.com/a/5046155/646444, this is not quite the same as asking if the wheels have both traveled the same distance — if the curve becomes tight enough that the radius of curvature is less than $L/2$, one of the wheels will be moving backwards during that time. If this backwards motion counts as positive term for “distance traveled”, then “distance traveled” will not be the same as “net forwards minus backwards motion”.
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The underlying math here is that $A$ defines a “holonomic constraint”, i.e., a set of relationships between derivatives of two subsets of a system’s configuration variables that is “integrable” into a functional relationship.
Note that this property does not hold for translational motion: Knowing the net roll angles of the wheels does not tell us how far the car has moved, and we would need the full history of the roll angles to reconstruct this motion. This gets into the idea of “nonholonomic constraints” that relate derivatives of subsets of a system’s variables, but cannot be turned into a functional relationship. (This topic requires a bit more involved discussion than seems appropriate here; the short form is that establishing that the exterior derivative of $A$ was zero was a special case of a more general check to see if a system is holonomic or nonholonomic.)